Over the next two years, the group toured and refined its lineup, adding multi-instrumentalist Gillian Lisee to the fold while embracing more elements of pop and experimental rock. In 2002, the Fruit Bats signed with Sub Pop and released their sophomore effort, Mouthfuls, the following spring. Two years later, having relocated to Seattle and expanded to a quartet, the band released Spelled in Bones. The album continued moving away from the folk-rock foundation of Echolocation, although elements of that rootsy sound remained.
Following the release of Spelled in Bones, Johnson took a break from the Fruit Bats to serve as a sideman for several other bands, including the Shins and Vetiver. He reconvened the group in 2008, having revised the lineup to include Christopher Sherman, Ron Lewis, Graeme Gibson, and Sam Wagster. The band returned to Chicago to record at Clava, the same studio that housed the sessions for Echolocation, and emerged with 2009's The Ruminant Band. He took a more solitary approach with Tripper, which appeared in 2011 and featured a pared down, story-based track list. -Heather Phares (AMG)